mohamed aly osman

Scopus Publications

Biological control of Tetranychus urticae mite by predatory mites compared to pesticides on strawberry plants under field conditions
Manal A. Abdel Razik, Zamzam M. Al Dhafar, Mohamed A. Osman, Mohamed E. Sweelam
Beni Suef University Journal of Basic and Applied Sciences, 2026
Background The two-spotted spider mite, Tetranychus urticae ( T. urticae ) is one of the most serious pests affecting strawberry plants. The use of various chemical pesticides has posed numerous environmental risks, necessitating the search for alternative solutions that are safer and more sustainable, such as the use of biological control agents, which offer an effective and environmentally friendly method in pest management. Methods This study was conducted to compare the biological control of the two-spotted spider mite by releasing predatory mites three times over two successive seasons, comparing to three pesticide systems applied by two methods as well as, to estimate the impact of the release processes and applied pesticides on strawberry yield. Results The predatory mites: Phytoseiulus persimilis ( P. persimilis ), and Neoseiulus californicus ( N. californicus ), demonstrated significant efficacy in reducing populations of T. urticae on strawberry plants during 2023 and 2024 seasons. P. persimilis outperformed the other species, recording the lowest numbers of eggs and motile stages of the pest, with reduction rates of 81.7% and 84.9% in the 2023 season, and 72.6% and 76.6% in the 2024 season. N. californicus ranked second in effectiveness, with reduction rates of 78.4% and 74.6% in the 2023 season, and 68.9% and 70.4% in the 2024 season. The chemical treatment of (thiamethoxam, micronized sulfur, and mineral oil) also showed good efficacy, particularly when applied as a foliar spray, with reduction rates of 85.7% and 81.5% in the 2023 season, and 68.1% and 71.4% in the 2024 season. Regarding strawberry yield, the biological treatment with P. persimilis achieved the highest increase in yield, with 38.7% in the 2023 season and 39.7% in the 2024 season. Among the chemical treatments, the foliar spray of thiamethoxam, micronized sulfur, and mineral oil resulted in the highest yield increase, with 46.2% in the 2023 season and 44% in the 2024 season. Conclusions It could be recommend the use of P. persimilis and N. californicus predators along with the treatment of (thiamethoxam-micronized sulfur-mineral oil) to control T. urticae infestation on strawberry plants under field conditions.
Nematicidial Activity of Five Cruciferous Seed Extracts in Controlling the Root-Knot Nematode Infecting Cucumber Plants
, S.M. Saadoon, S.B. Gad, , M.A. Osman, , M.I. Sergany, and
Arab Journal of Plant Protection, 2025
A greenhouse experiment was conducted to evaluate the effectiveness of five plant aqueous extracts of watercress, radish, turnip, cauliflower, and cabbage, either as seed extract or geminated seed extract, compared with oxamyl nematicide to protect cucumber plants from infection with the root knot nematode Meloidogyne incognita. In general, all the tested aqueous plant extracts and the chemical nematicide led to a significant decrease in the number of nematode root galls, egg masses, and eggs per gram of root as well as the number of larvae per 250 g of soil compared to the untreated inoculated control. The aqueous extracts of the germinated cauliflower seed and watercress seed had the highest nematicidal effect against root gall formation, egg-masses, and the number of eggs per gram of root. The cauliflower seed extract had the maximum nematicidal effect against nematode juveniles in soil, whereas the radish seed extract has turned the least effective. The tested aqueous plant extracts significantly increased the shoot’s fresh weight compared to the untreated inoculated control. Soil drenching with the germinated cruciferous seed extracts exhibited similar activity against M. incognita in terms of a significant increase in ascorbic acid, salicylic acid, and phenol content. Keywords: Plant extracts, nematicidal activity, Meloidogyne incognita, cucumber.
Effects of organic, non organic fertilizers, and selected novel pesticides on survival of two Entomopathogenic nematodes
Z. M. Al Dhafar, M. A. Abdel Razik, M. A. Osman, M. E. Sweelam
Brazilian Journal of Biology, 2025
In this study, the effect of six macro fertilizers i.e. urea, ammonium sulfate, ammonium nitrate, NPK, NP & DAP, and three micro elements i.e. Zn, Mn, and Fe at different concentrations as well as eight novel pesticides, thiamethoxam, spinetoram, azoxystrobin, chlorfenpyr, chlorantraniliprole, novaluron, lambda-cyhalothrin and avaunt at field recommended rate on two entomopathogenic nematodes (EPNs): Steinernema carpocapsae and Heterorhabditis bacteriophora were determined under laboratory conditions. The obtained results reported that S. carpocapsae was more tolerant to NPK; NP and DAP macro fertilizers than H. bacteriophora, showed the lowest mortality percentages (12.63, 9.47and12.63%), (15.79, 13.68 and 31.58%) and (20.21, 30.85 and 34.04%) at three tested concentrations (1g/liter water, 5g/liter water and10g/liter water), respectively. All tested micro fertilizers (Zn, Mn, and Fe) were lethal for two EPNs and induced 94.85, 100, 100 mortality % after five days of exposure. The morality percent increased as the concentration of fertilizers increased. With respect to tested pesticides, the mortality percentages of two EPN were ranged between 2.67-23.67% for S. carpocapsae and 3.33-18.33% for H. bacteriophora after five days of exposure. The tested macro fertilizers such as DAP, NP and NPK can be used safely at tested concentrations (5-10 g/ liter water) with Steinernema carpocapsae. All tested macro fertilizers cannot be used with Heterorhabditis bacteriophora. All micro fertilizers could be used with two EPNs. All tested pesticides can be successfully used for integrated plant protection systems. EPNs are tolerant to the tested pesticides and the tank-mix application is possible in most compounds except Lambda cyhalothrin which significantly reduced the virulence of tested nematodes.
Toxicity and biochemical effects of four pesticides on honey bee, Apis mellifera under laboratory conditions
Z. M. Al Dhafar, M. A. Abdel Razik, M. A. Osman, M. E. Sweelam
Brazilian Journal of Biology, 2025
The widespread use of insecticides can cause negative side effects on pollinators, resulting undesirable effects in crop productivity. So, this study was conducted to evaluate the toxicity and biochemical changes in honey bees, Apis mellifera (Hymenoptera: Apidae) enzymes, treated with four insecticides i.e. lambda-cyhalothrin, hexythiazox, fenpyroximate and thiamethoxam under laboratory conditions. The obtained results revealed that thiamethoxam was extremely toxic to A. mellifera adults (LC50=0.006 ppm) followed by lambda-cyhalothrin (LC50=0.053 ppm) and fenpyroximate (LC50=2.29 ppm) after 24 h of treatment, however, hexythiazox was relatively less toxic to bees (110.09 ppm). The sub-lethal concentration LC25 and LC50 of tested pesticides, reduced the activity of AChE, GST, MFO and esterase’s activities in honey bee adults, where, lambda-cyhalothrin and thiamethoxam induced the highest effects compared with other two pesticides and control. In contrast, the tested insecticides activated PPO enzyme at LC25 concentration, while PPO activity was reduced after treating adult bees with LC50. Lambda-cyhalothrin, thiamethoxam, fenpyroximate and hexythiazox disrupted the physiology of honey bees, thereby reducing the efficiency of this beneficial pollinator. Overall, the obtained results are valuable not only in evaluating the toxicity of common insecticides onto honey bees, but also in highlighting the validity of enzyme activities as appropriate indicators for exposure to agrochemicals.
Virulent Beauveria bassiana OR500626 strain effectively control Tetranychus urticae
J. K. Al-Zahrani, M. A. Osman, L. A. Aldakheel, A. K. Aldosary, S. Akhtar, H. Chakroun, M. A. Al Mustafa, M. A. Almuhawish, A. H. Al-Abdalall
Brazilian Journal of Biology, 2025
The entomopathogenic fungus Beauveria bassiana OR500626 plays a pivotal role as a biological control agent for various agricultural pests, including the two-spotted red spider mite, Tetranychus urticae Koch. In the current study, we tested the ability of a promising strain of the fungus isolated from the tissues of the red palm weevil, Rhynchophorus ferrugineus in the eastern region of the Kingdom of Saudi Arabia. B. bassiana virulence was estimated based on LC50 values calculated by Probit analysis of T. urticae, which were treated with a series of concentrations (1 × 102, 1 × 104, 1 × 106, and 1 × 108 conidia/ml). The mortality percentages of immature stages were 15.98-100%, males 15.94%-100%, and females (28.92%-100%) with an estimated LC50 value of 24.09×103, 35.43×103, and 11.03×103 conidia/ml, respectively. Furthermore, electron microscopy confirmed the ability of B. bassiana to infect T. urticae. The results indicated that B. bassiana spores attached to the T. urticae spider body within about 12 hours. The germination of spores increased within 24-36 and penetrated the T. urticae cuticle after 96 hours of exposure.
Acaricidal and insecticidal activities of entomopathogenic nematodes combined with rosemary essential oil and bacterium-synthesized silver nanoparticles against camel tick, Hyalomma dromedarii and wax moth, Galleria mellonella
Bander Albogami, Hadeer Darwish, Akram Alghamdi, Ahmed BahaaEldin Darwish, Wafa Mohammed Al-Otaibi, Mohamed A. Osman, Zamzam M. Al Dhafar, Abeer Mousa Alkhaibari, Abadi M. Mashlawi, Fadi Baakdah, Ahmed Noureldeen
Peerj, 2025
An innovative approach to ticks and insect pests management is necessary to mitigate the challenges posed by the indiscriminate use of chemical pesticides, which can lead to resistance development and environmental pollution. Despite their great potential, biological control agents have significant manufacturing, application, and stability limitations. Currently, using phytochemicals, biosynthesized nanoparticles, and bioagents to get rid of arthropods might be a good alternative that would make farmers less worried about residues and resistance. The present investigation was carried out to determine for the first time the in vitro acaricidal and insecticidal efficacies of endogenous two entomopathogenic nematodes (EPNs), Heterorhabditis indica and Steinernema sp. combined with either Proteus mirabilis-synthesized silver nanoparticles or Rosmarinus officinalis essential oil against the camel tick, Hyalomma dromedarii larvae and females, and greater wax moth, Galleria mellonella larvae as well. We also determined the potential effects of these treatments on the biological characteristics of H. dromedarii’s engorged females. We further investigated R. officinalis essential oil (EO) profiling and nanoparticle (AgNPs) characterization. All the evaluated combinations demonstrated synergistic effects on the larvae of G. mellonella and H. dromedarii, as well as on engorged females. When H. indica was mixed with EO or AgNPs, it worked well than when Steinernema sp. was mixed with EO or AgNPs. This was shown by the highest number of tick and insect mortalities and the lowest lethal concentration (LC50) values. One day after G. mellonella was exposed to H. indica (1,000 infective juveniles (IJs)) together with EO at 60 or 40 mg/mL, all tested individuals died. We obtained the same results when H. dromedarii females exposed to the same level of EPN with 60 mg/mL EO, and when H. dromedarii larvae treated with H. indica at 500 IJs + EO at 25 mg/mL. Treatments altered all biological parameters of engorged females, revealing extremely noticeable differences between the treated and untreated groups. Gas chromatography–mass spectrometry (GC-MS) analysis identified a total of 28 compounds in the R. officinalis EO. Visual observation showed a color change from yellow to dark brown for AgNPs biosynthesized from P. mirabilis; the transmission electron microscopy (TEM) image and ultraviolet–visible (UV-Vis) spectrum showed well-dispersed particles with a diameter of 5–45 nm; and the greatest surface plasmon peaked at 320 nm. The results demonstrated the high efficacy of combining EPN, H. indica, with EO to control tick and insect pests. This is due to its acaricidal activity on different stages of H. dromedarii, including larvae and engorged females, and its larvicidal effect on G. mellonella.
Efficacy of selected pesticides on Citrus Brown Mite, Eutetranychus orientalis (Acari: Tetranychidae) and the side effects on three predatory mites under citrus orchard conditions
Z. M. Al Dhafar, M. A. A. Abdel Razik, M. A. Osman, M. E. Sweelam
Brazilian Journal of Biology, 2024
The present study has been conducted to evaluate the effect of two sprays of seven pesticides at recommended dose on citrus brown mite, Eutetranychus orientalis and the side effects on their predatory mites, Euseius scutalis, Amblyseius swirskii, Phytoseiulus persimilis (Acari: Phytoseiidae) under field conditions at 2022 & 2023 seasons. The obtained results show that, all tested pesticides achieved high reduction % of E. orientalis ranged between (82.1-90.0%) and (81.6-87.1%) after the 1st and 2nd sprays of 2022 season, where it ranged between (84.9- 88.7%) and ( 79.7- 88.7%) after 1st and 2nd sprays of 2023 season. Abamectin recorded the highest reduction % against the citrus brown mite, whereas Congest pesticide recorded the lowest reduction % after the two sprays along 2022 & 2023 seasons. As for the side effects of tested pesticides on associated predatory mites, all pesticides were safely for E. scutalis numbers recording decrease % between (18.4-28.6%) and (16.2 -26.1%) after the 1st and 2nd spray at 2022 season , where it ranged between (15.3- 29.1%) and (19.6-32.0%) after the 1st and 2nd sprays of 2023 season. On contrary, imidacloprid was unsafely for E. scutalis numbers recording the highest mean decrease % after 1st and 2nd sprays during the two seasons. Also, all tested pesticides were safely for A. swirskii numbers, after the 1st and 2nd sprays of the two seasons recording decrease (from 10.9 to 28.1%) & (24.4 to 31.4%) for the 2022 season, and (19-38.9%) & (18.7-39.4%) at 2023 season. On contrary, imidacloprid was unsafely for A. swirskii numbers recorded the highest decrease % after 1st and 2nd sprays during the two seasons. As for, Ph. Persimilis numbers, all tested pesticides were safely, where it recorded low decrease % ranged between (17-33.8%) & (20.4-34.8%) after the 1st and 2nd sprays of 2022 season, and (24.3-39%) & (20.2-28.9%) after the 1st and 2nd sprays of 2023 season. On the other side, imidacloprid was unsafely for Ph. persimilis numbers recording the highest decrease % after the 1st and 2nd sprays during the two seasons. The present study proved that all tested pesticides were high effective against E. orientalis and appeared to be safely and selective for associated predatory mites except imidacloprid which was very harmful for all tested predatory mites, and it could be concluded that the tested pesticides, Fenpyroximate, Hexythiazox , Congest , Spirodiclofen, Abamectin, and Chlorfenapyr could be used in the Integrated Pest Management (IPM) programs for E. orientalis at citrus orchards.
Efficacy of novel insecticides against piercing sucking insects and their natural enemies on sweet pepper plants under field conditions
Z. M. Al Dhafar, M. A. A. Abdel Razik, M. A. Osman, M. E. Sweelam, N. O. Swelam
Brazilian Journal of Biology, 2024
Piercing sucking pests attacking sweet pepper plants cause significant losses to its yield. Considering the undesirable effects of synthetic pesticides, field studies were conducted to evaluate the impact of new pesticides against piercing sucking insect pests of sweet pepper, as well as, their effects on some predators and pepper yield along two seasons of 2021-2022. The obtained results indicated that all tested pesticides effectively suppressed the sucking insect populations (aphids, white fly, thrips) 1,7,14 and 21 days after treatment along two sprays during two seasons. Imidacloprid proved to be the superior one over all other treatments where it recorded mean reduction% (98.91 and 97.27%) & (94.8 and 95.19%), (86.23 and 76.64%) & (80.92 and 88.55%) and (77.68 and 78.44%) & (90.70 and 68.57%) in white fly, aphids and thrips, respectively at 1st and 2nd sprays at 2021 and 2022 seasons, respectively. As for side effects of tested insecticides on natural enemies, Dimethoate induced the highest decrease (60.85 and 69.33%) & (54.02 and 63.41%), (65.52 and 64.74%) & (59.23 and 58.38%) and (64.24 and 59.48%) & (61.66 and 60.8%) on Chrysoperla carnea, Paederus alfierii and Coccinella spp at 1st and 2nd sprays at 2021 and 2022 seasons, respectively. On contrary, Spintoram induced the lowest effects on Chrysoperla carnea, Paederus alfierii and Coccinella spp, recording decrease percent (25.41 and 19.84%) & (15.02 and 12.50%), (11.94 and 11.24%) (16.99 and 18.02%) and (18.73 and15.07%) & (18.35 and18.38%) at1st and 2nd sprays at 2021 and 2022 seasons, respectively. With respect to the effect of tested insecticides on pepper yield, all tested insecticides increased the yield of green pepper fruits compared with control. Imidacloprid achieved the highest fruit yields along two seasons 6.43 and 6.52 (ton / fed.4200 m2) with increase percent 34.53 and 36.04% in yield over control at 2021 and 2022 seasons, respectively.
Entomopathogenic fungi and their biological control of Tetranychus urticae: Two-spotted spider mites
Jawaher Khamis Al-Zahrani, Amira Hassan Al-Abdalall, Mohamed Aly Osman, Lena A. Aldakheel, Nada Faisal AlAhmady, Sumayh A. Aldakeel, Sayed AbdulAzeez, J. Francis Borgio, Medhat A. ElNaggar, Nadiyah M. Alabdallah, Mona M. Almustafa
Journal of King Saud University Science, 2023
The two-spotted spider mite (TSSM) Tetranychus urticae, is regarded as one of the most dangerous pests responsible for great losses in most of agricultural crops. It is a persistent pest in Saudi Arabia, especially in greenhouses where T. urticae is primarily controlled by chemical pesticides. The main problem for the two-spotted spider mites is its high resistance to pesticides and high fertility rate. In the long term, chemical pesticides cause health problems and economic losses, so it was necessary to search for a safe alternative method for human health and the environment. One of these alternative methods was the selection of plant varieties resistant to the TSSM, in addition to biological control that includes mites or predatory insects and entomopathogenic fungi. The growth, reproduction, and life-table parameters of T. urticae were examined in a laboratory setting with a 16L:8D photoperiod at 28±1°C and 65±5% RH, in the presence of three major members of Family: Solanaceae tomato, eggplant, and pepper. Pepper was shown to be less conducive to T. urticae growth and reproduction compared to eggplant and tomato. Tetranychus urticae proceeded through all five stages of its life cycle (egg, larva, protonymph, deutonymph, and adult) on tested solanaceous plants, and these plants significantly influenced its growth, reproduction, and Life-table parameters. Additionally, entomopathogenic fungi have been used against insects that have proven highly effective in controlling and reducing the density of two-spotted spider mites. Eight fungi were isolated from 80 insect and mite samples collected from several Saudi Arabia regions. Analysis of the 18S rRNA sequences revealed that the fungal strains identified as Beauveria bassiana, Fusarium sp. F. equiseti, F. oxysporium, Scopulariopsis brevicaulis1, S. brevicaulis2, Aspergillus sclerotiorum , and Penicillium citrinum. The ability of isolated fungi to secrete enzymes degrading the two-spotted spider mite cuticle, namely lipase, protease, and chitinase, were studied.
Planning for Future Jobs in Light of the Unified Saudi Classification of Educational Levels and Specializations—A Case Study of Graduate Students at Imam Abdul Rahman bin Faisal University
Ahmed Osman Ibrahim Ahmed, Anas Satti Satti Mohammed, Osman Saad Shidwan, Mohamednour Eltathir Ahmed Abdelgadir, Manal Mohamed EL Mekebbaty, Awad Mohamed Osman
Sustainability Switzerland, 2023
This study deals with the issue of planning for future jobs in light of the Unified Saudi Classification of Educational Levels and Specializations. We aimed to identify the mechanism used by graduates to choose a future job and to shed light on the Unified Saudi Classification of Educational Levels and Specializations. The problem addressed in this study is the identification of the optimal formula such that the graduate can benefit from this classification. The community studied is made up of students at the College of Applied Studies and Community Service at Imam Abdul Rahman bin Faisal University in Dammam in the period from 2019 to 2022. The sample included 129 male and female students, representing 20% of the research community. The selection was random, taking into account the homogeneity of the research community. We attempted to verify the validity of the hypothesis, stating that there is a statistically significant relationship between graduates’ preferences for their future jobs and their knowledge, represented by The Saudi Standard Classification of Scientific Levels and Specializations. A number of findings resulted from this study, most notably that there was a discrepancy regarding students’ preferences for future jobs based on their gender. We conclude with a number of recommendations, including the need to shed more light on the Unified Classification of Educational Levels and Specializations in Saudi Arabia and increase communication between scientific departments and employers.
Determination of thiamethoxam residues and dissipation kinetic in tomato plants and its efficacy against Bemisia tabaci under open field eco system
Z. M. Al Dhafar, M. A. A. Abdel razik, M. A. Osman, M. E. Sweelam
Brazilian Journal of Biology, 2023
Dissipation and Residues of Imidacloprid and Its Efficacy against Whitefly, Bemisia tabaci, in Tomato Plants under Field Conditions
Manal A. A. Abdel razik, Zamzam M. Al Dhafar, Aisha M. Alqahtani, Mohamed A. Osman, Mohamed E. Sweelam
Molecules, 2022
Combined use of Aqueous Plant Extracts for Controlling Meloidogyne incognita and Modulating Chemical Constituents in Tomato under Greenhouse Conditions
Doaa Khairy, Mohamed Ali Osman, Fatma Abdel Mohsen Mostafa
Pakistan Journal of Nematology, 2022
Managing gut microbiota through in ovo nutrition influences early-life programming in broiler chickens
Abdelrazeq M. Shehata, Vinod K. Paswan, Youssef A. Attia, Abdel-Moneim Eid Abdel-Moneim, Mohammed Sh. Abougabal, Mohamed Sharaf, Reda Elmazoudy, Wejdan T. Alghafari, Mohamed A. Osman, Mayada R. Farag, Mahmoud Alagawany
Animals, 2021
Molecular characterization and phylogenetic analysis of Rhynchophorus ferrugineus (Olivier) in Eastern Province, Saudi Arabia
Hussein Sabit, Shaimaa Abdel-Ghany, Zamzam Al-Dhafar, Osama A. Said, Jawad Ali Al-Saeed, Yousef Ahmed Alfehaid, Mohamed Aly Osman
Saudi Journal of Biological Sciences, 2021
Biological responses of the two-spotted spider mite, Tetranychus urticae to different host plant
Mohamed Aly Osman, Zamzam M. Al Dhafar, Aisha M. Alqahtani
Archives of Phytopathology and Plant Protection, 2019
Development and reproductive potential of Tyrophagus putrescentiae (Acari: Acaridae) on plant-parasitic nematodes and artificial diets
Doaa Abd El-Maksoud Abou El-Atta, Mohamed Ali Osman
Experimental and Applied Acarology, 2016
Life table parameters of the predatory mite, Cydnoseius negevi (Swirski & amitai) affected by prey density of Tetranychus urticae koch (acari: Phytoseiidae, tetranychidae)
Egyptian Journal of Biological Pest Control, 2016
Potential use of stigmaeid mites for biological control of orchard pests
Egyptian Journal of Biological Pest Control, 2015
Effects of temperature on the life-history traits of Sancassania (Caloglyphus) berlesei (Acari: Astigmatina: Acaridae) feeding on root-knot nematodes, Meloidogyne spp. (Nematoda: Meloidogynidae)
Doaa Abd El-Maksoud Abou El-Atta, Noureldin Abuelfadl Ghazy, Mohamed Ali Osman
Experimental and Applied Acarology, 2014
Functional response of the phytoseiid mite Typhlodromus negevi Swirski and Amitai to the two-spotted spider mite Tetranychus urticae (Acari: Tetranychidae)
Aly H. Rasmy, G.M. Abou-Elella, M.A. Osman
Archives of Phytopathology and Plant Protection, 2014
Host plant mediates foraging behavior and mutual interference among adult Stethorus gilvifrons (Coleoptera: Coccinellidae) preying on Tetranychus urticae (Acari: Tetranychidae)
Mohamed H. Bayoumy, Mohamed A. Osman, J. P. Michaud
Environmental Entomology, 2014
Biology and life tables of Esieus scutalis (A.-H.) reared on different kinds of food
A.H. Fouly, O.A. Nassar, M.A. Osman
Journal of Entomology, 2013
Thermal requirements and biological life table parameters of the predatory mite Amblyseiella denmarki (Zaher and El-Borolossy) (Acari: Phytoseiidae)
G.M. Abou-Elella, M.A. Osman, E.M. El-Saidey
Archives of Phytopathology and Plant Protection, 2012
Functional response of Orius albidipennis (Hemiptera: Anthocoridae) to the two-spotted spider mite Tetranychus urticae (Acari: Tetranychidae)
Nesrin A. El-Basha, M.S. Salman, M.A. Osman
Journal of Entomology, 2012
Effect of prey stages of the two-spotted mite Tetranychus urticae on functional response of the coccinellid predator Stethorus gilvifrons
M. Osman, M. Bayoumy
Acta Phytopathologica Et Entomologica Hungarica, 2011
Temperature influence on biology, thermal requirement and life table of the predatory mites Agistemus exsertus Gonzalez and Phytoseius plumifer (Can. & Fanz.) reared on Tetranychus urticae Koch
Aly H. Rasmy, M. A. Osman, G.M. Abou-Elella
Archives of Phytopathology and Plant Protection, 2011

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